San Diego State University Herbarium (SDSU)


Publications utilizing and citing specimens from the SDSU Herbarium

2026
Simpson, M. G., M. E. Mabry, P. D. Blischak, T. Böhnert, C. M. Guilliams, K. E. Hasenstab-Lehman, F. Luebert, N. García, Á. Castañeda, M. Weigend,

     and R. A. Scherson. 2026. Resolving the taxonomic enigma of Nesocaryum stylosum (Boraginaceae): phylogenetic evidence for its reclassification
     as Cryptantha stylosa. PhytoKeys 269:141-169. https://doi.org/10.3897/phytokeys.269.176263

2025
Rebman, J. P., J. Gregory, and M. G. Simpson. 2025. Checklist of the Vascular Plants of San Diego County. 6th ed. San Diego Natural History Museum,
     San Diego, California. https://sdplantatlas.org/pdffiles/Rebman_etal2025-ChecklistSDCo.pdf

2024
Simpson, M. G. and D. A. York. 2024. Cryptantha whippleae (Boraginaceae), a new serpentine-adapted species endemic to northern California, U.S.A.
      PhytoKeys 247: 155–172. https://doi.org/10.3897/phytokeys.247.132060
Simpson, M. G., L. M. Simpson, and J. André. 2024. Cryptantha acrimuricata (Boraginaceae), a distinctive new taxon of series Muricatae.
      Phytokeys 250: 193–213. https://doi.org/10.3897/phytokeys.250.1386352023

2023
Simpson, M. G. and L. Gross. 2023. Noteworthy Collection: Cryptantha crassisepala (Torrey & A.Gray) Greene var. elachantha I.M.Johnston.
      Madroño 70:4–8. https://doi.org/10.3120/0024-9637-70.1.4

2022
Simpson, M. G., R. Vanderhoff, J. Bailey, F. Roberts, M. Mulligan, J. P. Rebman, N. Walsh, and M. Koekemoer. 2023. Taxonomic identity of a
      recently naturalized Senecio species in California. Madroño 69:286–298. https://doi.org/10.3120/0024-9637-69.3.286
Moroni, P. and M. G. Simpson. 2022. Revisión taxonómica del genero Johnstonella (Boraginaceae s. str.) en la Argentina. Darwiniana Nueva
      Serie 10:260–270. http://www.ojs.darwin.edu.ar/index.php/darwiniana/article/view/1046
Rebman, J. P. and M. G. Simpson. 2022. Three new species of Cryptantha (Boraginaceae) from the Southern Channel Islands of California.
      Madroño 69: 111-134. https://doi.org/10.3120/0024-9637-69.1.111
Simpson, M. G., K. Hasenstab-Lehman, M. Mabry, and M. Muñoz. 2022. Johnstonella punensis (Boraginaceae), a new species endemic to the
      dry Puna of Chile. PhytoKeys 197:149–164. https://doi.org/10.3897/phytokeys.197.84833

2021
Simpson, M. G. and J. P. Rebman. 2021. A new species of Cryptantha restricted to dunes in northwestern Baja California, Mexico.
      Madroño 68:127-137. https://doi.org/10.3120/0024-9637-68.2.127
Simpson, M. G. and J. P. Rebman. 2021. Research in Boraginaceae: A new variety of Cryptantha maritima, Cryptantha pondii resurrected,
      and Johnstonella echinosepala transferred back to Cryptantha. Phytotaxa 509:185–210. https://doi.org/10.11646/phytotaxa.509.2.3
Yost, J. M., K. D. Pearson, J. Alexander, E. Gilbert, L. A. Hains, T. Barry, R. Bencie, P. Bowler, B. Carter, R. E. Crowe, E. Dean, J. Der, A. Fisher, K.
      Fisher, L. Flores-Renteria, C. M. Guilliams, C. Hatfield, L. Hendrickson, T. Huggins, L. Janeway, C. Lay, A. Litt, S. Markos, S. J. Mazer, D. Mccamish,
      L. Mcdade, M. Mesler, B. Mishler, M. Nazaire, J. Rebman, L. Rosengreen, P. W. Rundel, D. Potter, A. Sanders, K. C. Seltmann, M. G. Simpson,
      G. A. Wahlert, K. Waselkov, K. Williams, and P. S. Wilson. 2020. The California Phenology Collections Network: Using digital images to investigate
      phenological change in a biodiversity hotspot. Madroño 66:130-141. https://doi.org/10.3120/0024-9637-66.4.130

2020
Simpson, M. G. and R. B. Kelley. 2020. Cryptantha. Pp. 394-403 in T. J. Stephen C. Meyers, Katie E. Mitchell, Tanya Harvey & Linda K. Hardison (ed.),
      Flora of Oregon. Volume 2: Dicots A-F. Brit Press, Botanical Research Institute of Texas, Fort Worth, Texas.
Simpson, M. G., J. Stephens, and S. Yang. 2020. Noteworthy Collections: Johnstonella angelica (I.M.Johnston) Hasenstab & M.G.Simpson.
      Madroño 67:5-8. https://doi.org/10.3120/0024-9637-67.1.5

2019
Benet-Pierce, N. and M. G. Simpson. 2019. The taxonomy of Chenopodium hians, C. incognitum, and ten new taxa within the narrow-leaved
      Chenopodium
group in western North America, with special attention to California. Madroño 66:56–75.
Simpson, M. G., M. E. Mabry, and K. Hasenstab-Lehman. 2019. Transfer of four species of Cryptantha to the genus Johnstonella (Boraginaceae).
      Phytotaxa 425:279–289. https://doi.org/10.11646/phytotaxa.425.5.3

2018
Mabry, M. E.* and M. G. Simpson. 2018. Evaluating the monophyly and biogeography of Cryptantha (Boraginaceae). Systematic Botany 43: 53-76.
    http://dx.doi.org/10.1600/036364418X696978

2017
Aerne, L. and M. G. Simpson. 2017. Vegetative anatomy of the Haemodoraceae and its phylogenetic significance.
     International Journal of Plant Sciences 178: 117-156.
Benet-Pierce, N. and M. G. Simpson. 2017. Taxonomic recovery of the species in the Chenopodium neomexicanum complex and description of
    Chenopodium sonorense
sp. nov. Journal of the Torrey Botanical Society 144: 339-356.
    http://www.bioone.org/doi/10.3159/TORREY-D-16-00013.1
Guilliams, M. C.*, K. Hasenstab-Lehman*, M. Mabry* and M. G. Simpson. 2017. Memoirs of a frequent flier: Phylogenomics reveals 18 long-distance
    dispersals between North America and South America in the popcorn flowers (Amsinckiinae, Boraginaceae).
    American Journal of Botany 104: 1717-1728. https://doi.org/10.3732/ajb.1700369
Simpson, M. G., C. M. Guilliams*, K. E. Hasenstab-Lehman*, M. E. Mabry*, and L. Ripma*. 2017. Phylogeny of the popcorn flowers: Use of genome
    skimming to evaluate monophyly and interrelationships in subtribe Amsinckiinae (Boraginaceae). Taxon 66: 1406–1420.
    https://doi.org/10.12705/666.8
Simpson, M. G. and R. B. Kelley. 2017. Cryptantha nevadensis var. rigida elevated to species with a new name, C. juniperensis.
    Phytotaxa 295: 227-236.

2016

Mabry, M. E.*, R. A. Dowdy*, L. M. Simpson, J. P. Rebman, and M. G. Simpson. 2016. Taxonomy of the winged popcorn flower: Cryptantha pterocarya
     
(Boraginaceae). Phytotaxa 253:97-130. (pdf)
McCabe, S. W., M. W. Dodero, and M. G. Simpson. 2016. Dudleya hendrixii A New, Rare Species From Colonet Mesa, Baja California.
    Madroño 63: 359-365.
Simpson, M. G., L. M. Simpson, and J. P. Rebman. 2016. A new, large-flowered variety of Eremocarya micrantha (Boraginaceae). Madroño 63: 39-54.

2014
Benet-Pierce, N. and M. G. Simpson. 2014. The taxonomy of Chenopodium desiccatum and C. nitens, sp. nov.
   Journal of the Torrey Botanical Society 141:161-172.
Rebman, J. P. and M. G. Simpson. 2014. Checklist of the Vascular Plants of San Diego County, 5th edition. San Diego Natural
     History Museum Publications. (link)

Ripma, L.*, M. G. Simpson, and K. Hasenstab-Lehman*. 2014. Geneious! Simplified genome skimming methods for
   phylogenetic systematic studies: A case study in Oreocarya (Boraginaceae). Applications in Plant Sciences 2: 1400062.
   doi: 10.3732/apps.1400062 (pdf).
Simpson, M. G., R. Dowdy*, J. P. Rebman, R. B. Kelley, and L. M. Simpson. 2014. The recognition of two species in
   Eremocarya (Boraginaceae): Evidence from fornix bodies, nutlets, corolla size, and biogeography. Madroño 61: 259–275.

2013
Guilliams, C. M.*, B. A. Veno, M. G. Simpson, and R. B. Kelley. 2013. Pectocarya anisocarpa, a new species of Boraginaceae,
     and a revised key for the genus in western North America. Aliso 31: 1-13.
Silveira, M.* and Simpson, M. G. 2013. Phylogenetic systematics of the mesa mints: Pogogyne (Lamiaceae).
     Systematic Botany 38: 792-794.
Simpson, M. G., J. P. Rebman, K. E. Hasenstab-Lehman, C. M. Guilliams, and P. McConnell. 2013. Cryptantha wigginsii
     
(Boraginaceae): A presumed extinct species rediscovered. Madroño 60(1): 24-34.
Simpson, M. G. and J. P. Rebman. 2013. A new species of Cryptantha (Boraginaceae) from the Sierra de San Pedro Mártir,
     Baja California, Mexico. Madroño 60(1): 35-45.
Yost, J. M., M. Bontrager, S. McCabe, D. Burton*, M. G. Simpson, K. M. Kay, and M. Ritter. 2013. Phylogenetic relationships
     and evolution in Dudleya (Crassulaceae). Systematic Botany 38: 1096-1104.

2012
Dodero, M. W.* and M. G. Simpson. 2012. Dudleya crassifolia (Crassulaceae), a new species from northern Baja California,
     Mexico. Madroño 59: 223-229.
Hasenstab-Lehman, Kristen E.* and M. G. Simpson. 2012. Cat’s Eyes and Popcorn Flowers: Phylogenetic Systematics of the
     Genus Cryptantha s.l. (Boraginaceae). Systematic Botany 37: 738–757.
Kelley, R. B., M. G. Simpson, and K. E. Hasenstab. 2012. Cryptantha. Pp. 455-468 in B. G. Baldwin, D. H. Goldman, D. J. Keil, R. Patterson,
     T. J. Rosatti, and D. H. Wilken (eds.), The Jepson Manual: Vascular Plants of California. Second Edition. University of California Press, Berkeley,
     Los Angeles, London.
Silveira, M., M. G. Simpson, and J. D. Jokerst. 2012. Pogogyne. Pp. 850-851 in B. G. Baldwin, D. H. Goldman, D. J. Keil, R. Patterson, T. J. Rosatti,
     and D. H. Wilken (eds.), The Jepson Manual: Vascular Plants of California. Second Edition. University of California Press, Berkeley, Los Angeles,
     London.

2011
Guilliams, C. M.*, Simpson, M. G., and J. P. Rebman. 2011. Calyptridium parryi var. martirense (Montiaceae), a new taxon endemic to the
     Sierra San Pedro Mártir, Baja California, Mexico. Madroño 58(4): 258-266.

2010
Benet-Pierce, N. and M. G. Simpson. 2010. Chenopodium littoreum (Chenopodiaceae): a new goosefoot from dunes of
     south-central coastal California. Madroño 57: 64–72.
Simpson, M. G., M. Silveira*, and C. M. Guilliams.* 2010. Taxonomy of Calyptridium parryi (Montiaceae). Madroño 57: 154-160.

2009
Simpson, Michael G. and Kristen E. Hasenstab*. 2009. Cryptantha of Southern California. Crossosoma 35: 1-59.
Pierce, Nuri and Michael G. Simpson. 2009. Polyaperturate pollen types and ratios of heteromorphism in the monocot genus
     Conostylis R. Br. (Haemodoraceae). Australian Systematic Botany 22: 1-15.

2003
Hargrove, Lori and Michael G. Simpson. 2003. Ultrastructure of heterocolpate pollen in Cryptantha (Boraginaceae).
      International Journal of Plant Sciences 164(1): 137-151.

1999
Marsden, Kim L.* and Michael G. Simpson. 1999. Eryngium pendletonensis (Apiaceae), a new species from southern California.
     Madroño 46(1): 55-58.

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